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|- | style="text-align:center;" | Fuji apple
Fuji apple |- style="text-align:center;" ! style="background: lightgreen;" | Scientific classification |- style="text-align:center;" |

|- valign=top |Kingdom:||Plantae |- valign=top |Division:||Magnoliophyta |- valign=top |Class:||Magnoliopsida |- valign=top |Order:||Rosales |- valign=top |Family:||Rosaceae |- valign=top |Subfamily:||Maloideae |- valign=top |Genus:||Malus |} |- style="text-align:center; background:lightgreen;" !Species |- | Malus domestica
Malus sieversii |}

Apple is the fruit (pome) of the genus Malus belonging to the family Rosaceae, and is one of the most widely cultivated tree fruits. Table apples are of the species M. domestica or hybrids of it.


Botanical origins

The wild ancestor of Malus domestica is Malus sieversii (which has no common name), a tree still found wild in the mountains of central Asia in southern Kazakhstan, Kyrgyzstan, Tajikistan, and Xinjiang Province, China. M. sieversii resists many diseases and pests that affect domestic apples, and research with it to develop new disease-resistant apples is continuing.

Other species that were previously thought to have made contributions to the genome of the domestic apples are Malus baccata and Malus sylvestris, but there is no hard evidence for this in older apple cultivars. These and other Malus species have been used in some recent breeding programmes to develop apples suitable for growing in climates unsuitable for M. domestica, mainly for increased cold tolerance.

Apples have been a very important food in all cooler climates, and is probably the earliest tree to be cultivated. To a greater degree than other tree fruit, except possibly citrus, apples store for months while still retaining much of their nutritive value. Winter apples, picked in late autumn and stored just above freezing, have been an important food in Asia and Europe for millennia, and the United States since the arrival of Europeans.

Apple cultivars

There are more than 7,500 known cultivars of apples. Different cultivars are available for temperate and subtropical climates. Apples do not flower in tropical climates because they have a chilling requirement.

Commercially-popular apple cultivars are soft but crisp. Other desired in modern commercial apple breeding are a colorful skin, absence of russeting, ease of shipping, lengthy storage ability, high yields, disease resistance, typical "Red Delicious" apple shape, long stem (to allow pesticides to penetrate the top of the fruit), and popular flavor.

Old cultivars are often oddly shaped, russeted, and have a variety of textures and colors. Many of them have excellent flavour (often better than most modern cultivars), but may have other problems which make them commercially unviable, such as low yield, liability to disease, or poor tolerance for storage or transport. Few old cultivars are still produced on a large scale, but many have been kept alive by home gardeners and farmers that sell directly to local markets. Many unusual and locally important cultivars with their own unique flavor and appearance are out there to discover; apple conservation campaigns have sprung up around the world to preserve such local heirlooms from extinction.

Although most cultivars are bred for eating fresh (dessert apples), some are cultivated specifically for cooking (culinary apples) or producing cider. Cider apples are typically too tart and astringent to eat fresh, but they give the beverage a rich flavor that dessert apples cannot.

The following is a list of common cultivars and where they are grown. The year and place of origin is also listed:

Flavors of apples

Modern apples are, as a rule, sweeter than older cultivars. Most North Americans and Europeans favor sweet, subacid apples, but tart apples have a strong minority following. Extremely sweet apples with barely any acid flavor are popular in Asia.

Tastes in apples vary from one person to another and have changed over time. As an example, the U.S. state of Washington made its reputation for apple growing on Red Delicious. In recent years, many American apple connoisseurs regard Red Delicious as inferior to varieties such as Fuji and Gala due to the mild flavor and soft texture of the Red Delicious.

Apple breeding

Apple tree in flower
Apple tree in flower

Like most perennial fruits, apples are ordinarily propagated asexually by grafting. Seedling apples differ from their parents, sometimes radically. Most new apple cultivars originate as seedlings, which either arise by chance or are bred by deliberately crossing cultivars with promising characteristics. The words seedling, pippin, and kernel in the name of an apple cultivar suggest that it originated as a seedling. Apples can also form bud sports (mutations on a single branch). Some bud sports turn out to be improved strains of the parent cultivar. Some differ sufficiently from the parent tree to be considered new cultivars.

Some breeders have crossed ordinary apples with crabapples or unusually hardy apples in order to produce hardier cultivars. For example, the Excelsior Experiment Station of the University of Minnesota has, since the 1930s, introduced a steady progression of important hardy apples that are widely grown, both commercially and by backyard orchardists, throughout Minnesota and Wisconsin. Its most important introductions have included Haralson (which is the most widely cultivated apple in Minnesota), Wealthy, Honeygold, and Honeycrisp. The sweetness and texture of Honeycrisp have been so popular with consumers that Minnesota orchards have been cutting down their established, productive trees to make room for it, a heretofore unheard of practice.

Growing apples

Apple orchard
Apple orchard

Starting an orchard

Apple orchards are established by planting two or three year old trees. These small trees are usually purchased from a nursery where they are produced by grafting or budding. First, a rootstock is produced either as a seedling or cloned using tissue culture or layering. This is allowed to grow for a year. Then, a small section of branch called a scion is obtained from a mature apple tree of the desired cultivar. The upper stem and branches of the rootstock are cut away and replaced with the scion. In time, the two sections grow together and produce a healthy tree.

Rootstocks affect the ultimate size of the tree. While many rootstocks are available to commercial grower, those sold to homeowners who want just a few trees are usually one of two cultivars: a standard seedling rootstock that gives a full-size tree, or a semi-dwarf rootstock that produces a somewhat smaller tree. Dwarf rootstocks are generally more susceptible to damage from wind and cold. Full dwarf trees are often supported of posts or trellises and planted in high density orchards which are much simpler to culture and greatly increase productivity per unit of land.

Some trees are produced with a dwarfing "interstem" between a standard rootstock and the tree, resulting in two grafts.

After the small tree is planted in the orchard, it must grow for 3-5 years (semi-dwarf) or 4-10 years (standard trees) before it will bear sizable amounts of fruit. Good training of limbs and careful nipping of buds growing in the wrong places, are extremely important during this time, to build a good scaffold that will later support a fruit load.


Apples are relatively indifferent to soil conditions and will grow in a wide range of pH values and fertility levels. They do require some protection from the wind and should not be planted in low areas that are prone to late spring frosts. Apples do require good drainage, and heavy soils or flat land should be tiled to make certain that the root systems are never in saturated soil.


Apples are self-incompatible and must be cross-pollinated. Pollination management is an important component of apple culture. Before planting, it is important to arrange for pollenizers, cultivars of apple or crab apple that provide plentiful, viable and compatible pollen. Orchard blocks may alternate rows of compatible cultivars, or may have periodic crab apple trees, or grafted-on limbs of crab apple. Some cultivars produce very little pollen, or the pollen is sterile, so these are not good pollenizers. Quality nurseries have pollenizer compatibility lists.

Growers with old orchard blocks of single cultivars sometimes provide bouquets of crab apple blossoms in drums or pails in the orchard for pollenizers. Home growers with a single tree, and no other cultivars in the neighborhood can do the same on a smaller scale.

During the flowering each season, apple growers usually provide pollinators to carry the pollen. Honeybee hives are most commonly used, and arrangements may be made with a commercial beekeeper who supplies hives for a fee. Orchard mason bees are also used as supplemental pollinators in commercial orchards. Home growers may find these more acceptable in suburban locations because they do not sting. Some wild bees such as carpenter bees and other solitary bee s may help. Bumble bee queens are sometimes present in orchards, but not usually in enough quantity to be significant pollinators.

Symptoms of inadequate pollination are small and misshapen apples, and slowness to ripen. Count the seeds to evaluate pollination. Well pollinated apples are the best quality, and will have 7 to 10 seeds. Less than 3 seeds will usually not mature and will drop from the trees in the early summer. Inadequate pollination can result from either a lack of pollinators or pollenizers, or from poor pollinating weather at flowering time. It generally requires multiple bee visits to deliver sufficient grains of pollen to accomplish complete pollination.

A common problem is a late frost that destroys the delicate outer structures of the flower. It is best to plant apples on a slope for air drainage, but not on a south facing slope (in the northern hemisphere) as this will encourage early flowering and increase susceptibilty to frost. If the frost is not too severe, the tree can be wetted with water spray before the morning sun hits the flowers, and it may save them. Frost damage can be evaluated 24 hours after the frost. If the pistil has turned black, the flower is ruined and will not produce fruit.

Growing apples near a body of water gives an advantage by slowing spring warm up, which retards flowering until frost is less likely. Areas of the USA, such as the eastern shore of Lake Michigan, the southern shore of Lake Ontario, and around some smaller lakes, where this cooling effect of water, combined with good, well-drained soils, has made apple growing concentrations possible in these areas.

Home growers may not have a body of water to help, but can utilize north slopes or other geographical features to retard spring flowering. Apples (or any fruit) planted on a south facing slope in the northern hemisphere (or north facing in the southern hemisphere), will flower early and be particularly vulnerable to spring frost.


Apples are prone to biennial bearing . If the fruit is not thinned when the tree carries a large crop, it may produce very little flower the following year. Good thinning helps even out the cycle, so that a reasonable crop can be grown every year.

Pests and diseases

The trees are susceptible to a number of fungal and bacterial diseases and insect pests. Nearly all commercial orchards pursue an aggressive program of chemical sprays to maintain high fruit quality, tree health, and high yields. A trend in orchard management is the use of Integrated Pest Management (IPM), which reduces needless spraying when pests are not present, or more likely, are being controlled by natural predators.

Spraying for insect pests must never be done during flowering because it kills pollinators. Nor should bee-attractive plants be allowed to establish in the orchard floor if insecticides are used. Dutch white clover is a component of many grass seed mixes, and many bees are poisoned by insecticides while visiting the flowers on the orchard floor.

Among the most serious disease problems are fireblight, a bacterial disease; and Gymnosporangium rust, apple scab , and black spot , three fungal diseases.

The plum curculio is the most serious insect pest. Others are Apple Maggot and codling moth .

Apples are difficult to grow organically, though a few orchards have done so with commercial success, using disease-resistant cultivars and the very best cultural controls. The latest tool in the organic repertoire is to spray a light coating of kaolin clay, which forms a physical barrier to some pests, and also helps prevent apple sun scald.


Mature trees typically bear 100-200 kg (5-10 bushels) of apples each year. Apples are harvested using three-point ladders that are designed to fit amongst the branches. A few cultivars, left unpruned, will grow to be extremely large, causing them to bear a great deal of fruit that is difficult to harvest. Dwarf trees will bear about 50-100 kg (3-5 bushels) of fruit per year.

Cultivars vary in their yield and the ultimate size of the tree, even when grown on the same rootstock.



45 million metric tons of apples were grown worldwide in 2002, with a value of about 10 billion USD. China produced almost half of this total. The United States is the second leading producer, accounting for 10% of world production. Turkey is also a leading producer. France, Italy, South Africa and Chile are among the leading apple exporters.

Today, more than 60% of all the apples sold commercially in the United States are grown in Washington state. This may change. Imported apples from New Zealand and other more temperate areas are competing with domestic production and increasing each year.


Apples can be canned, juiced, and optionally fermented to produce apple juice, cider, vinegar, and pectin. Distilled apple cider produces the spirits applejack and Calvados.

Apples are an important ingredient in many winter desserts, for example apple pie, apple crumble and apple cake. They are often eaten baked or stewed , and they can also be dried and eaten or re-consitituted (soaked in water, alcohol or some other liquid) for later use. Puréed apples are generally known as apple sauce. Apples are also made into apple butter and apple jelly. They are also used cooked in meat dishes.

Health benefits

Apples have been proven to reduce the risk of colon cancer, prostate cancer and lung cancer. They can also help with heart disease, weight loss and controlling cholesterol.

Cultural aspects

The ancient Kazakh city of Almaty, 'Father of Apples' (Kazakh language alma, apple, + ata, father), owes its name to the forests of wild apples (Malus sieversii) found naturally in the area.

Apples were very important in many ancient cultures, including Norse, Roman and Greek beliefs. See Pleiades and Idun for examples.

In Norse myth, Idun was the keeper of the 'apples of immortality' which kept the Gods young. The 'fruit-bearing tree' referred to by Tacitus in his description of Norse runic divination may have been the apple.

Although the "forbidden fruit" in the book of Genesis is not identified, popular European Christian tradition has held that it was an apple that Adam and Eve ate. This tradition was probably solidified by artistic renderings of the fall from Eden featuring an apple; some kind of fruit had to be pictured and the apple was probably the most familiar fruit to the artists. The larynx in the human throat has been called Adam's apple because of a notion that it was caused by the forbidden fruit sticking in the throat of Adam.

The name of the Russian party Yabloko means "apple". Its logo represents an apple in the constructivist style.
The name of the Russian party Yabloko means "apple". Its logo represents an apple in the constructivist style.

Apples are eaten with honey at the Jewish New Year of Rosh Hashanah to symbolize a sweet new year.

The apple flower is the state flower of Arkansas.

In some cultures, the apple is a symbol of immortality, love, or sexuality. The Greek hero Heracles had to find the Hesperides' golden apples as one of his Twelve Labours. Another Greek mythological figure, Paris, had to give a golden apple inscibed Kallisti – "For the most beautiful one", or "To the Prettiest One" – (which had come from the goddess of discord, Eris) to the most beautiful goddess, thus indirectly causing the Trojan War. Atalanta, also of Greek mythology, was distracted during a race by three golden apples thrown for that purpose by a suitor, Hippomenes. In ancient Greece, throwing an apple at a person's bed was an invitation for Censored page. Celtic mythology includes a story about Conle who receives an apple which feeds him for a year but also makes him irresistibly desire fairyland. Another story claims that if an apple is peeled into one continuous ribbon and thrown behind a woman's shoulder, it will land in the shape of the future husband's initials. Danish folklore says that apples wither around adulterers.

In some places, dunking for apples is a traditional Halloween activity. Apples are said to increase a woman's chances of conception as well as remove birthmarks when rubbed on the skin. They are commonly considered healthy, leading to the proverb an apple a day keeps the doctor away. In the United States, Denmark and Sweden, an apple is a traditional gift for a teacher.

In the UK, a toffee apple is a traditional confection made by coating an apple in hot toffee and allowing it to cool. Similar treats in the US are candy apples (coated in a hard shell of crystallized sugar syrup), and caramel apples , coated with cooled caramel.

See also

External links and references

  • Over 2000 apples in the UK's National Fruit Collections
  • U.S Apple Association Guide with some years and places of cultivar origins
  • Apple Facts from the UK's Institute of Food Research
  • Washington State Apple Commission
  • Wild apples in Kazakhstan: 1995 and 1996 expeditions

Last updated: 02-02-2005 10:10:25
Last updated: 03-13-2005 10:41:26